I wouldn't think there'd be that much between towing/power cababilites of either as they are very similar drive train and rated at the same towing capacity, and given the difference in weight of the vehile, the gap is closed by the ratio and number of cogs to play with (6spd vs 5sd).
But I reckon you will notice the ride and comfort a hellava lot though when not hitched up - I was tossing up between a Prado and Hilux in 2010 and opted for the Navara based on cost and it was more passenger car like than the Hilux. I drive Prados and Hiluxs for work (not towing though) and know exactly which one I’d prefer to drive on the long haul...
There’s lot to consider – family vs. two people (rear leg room), what goes in the back when on tour and not on tour, how much towing will you be doing… A big winner for the Prado in my book is the dual fuel tank for maximum range between pit stops when towing.
Decisions decisions… just wish it was me with this delima J